Gulf of Maine Council on the Marine Environment

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Identification of Important Habitats in Coastal New Hampshire

Chapter 9. American Shad

American shad (Alosa sapidissima) is an anadromous clupeid fish, ranging from Florida to the St Lawrence River. It formerly was abundant enough to be of major importance as a food source; dams and degradation of water quality in spawning rivers have greatly reduced historic runs. We mapped shad habitat using a combination of occurrence (known use) information and habitat relationships to environmental features.

SPAWNING HABITAT

Shad ascend freshwater tributaries and spawn in slow-flowing sections of rivers (Scott and Scott 1988). Spawning occurs at water temperatures < 23 degrees C. (Scott and Scott 1988) and > 10 degrees C. (Leim 1924, Williams and Daborn 1984 cited in Scott and Scott 1988). Shad spawn in Canadian tributaries between May and July (Scott and Scott 1988). Because of the paucity of other information on spawning habitat requirements of American shad, we mapped spawning habitat only from known occurrences (D. Grout, New Hampshire Fish and Game Department). These areas were scored as 10 (0 - 10 scale) since they in fact must meet shad environmental requirements (Figure of Spawning Habitat).

LARVAL HABITAT

Shad eggs are pelagic and are carried downstream by the current (Weiss-Glanz et al. 1988).

Incubation time of eggs is dependent on water temperature (Scott and Crossman 1973, Marcy 1976). Eggs cease developing at water temperatures of 7 degrees C. and abnormalities occur at 22 degrees C. (Leim 1924). Eggs and larvae survive in water at salinities between 7.5 and 15 ppt, but mortality occurs when salinity reaches 22.5 ppt (Leim 1924). The salinity regime in which eggs hatch successfully remains unclear (Weiss-Glanz et al. 1988). At hatching, larvae are approximately 7 mm total length (TL) and are planktonic (Marcy 1976), and grow to 12 mm TL when the yolk-sac is absorbed (Jones et al. 1978). Transformation to the juvenile stage occurs at 25 - 28 mm TL at 2 - 3 weeks of age (Jones et al. 1978).

Distribution of larval habitat was mapped only within rivers where spawning is known to occur. Habitat was regarded as those reaches having salinities < 15 ppt during the spring season (April - June). These were given a score of 7.5 (0 - 10 scale) since actual presence of larvae has not been documented (Figure of Larval Habitat). The paucity of information on other environmental requirements prevented development of a more rigorous model.

JUVENILE HABITAT

Juveniles occur in natal rivers during summer (Weiss-Glanz et al. 1988). Decreasing water temperature is the stimulus for downstream movement of juveniles into brackish water and finally to the sea (Weiss-Glanz et al. 1988). Because of the paucity of information on juvenile habitat requirements, we mapped early juvenile habitat as only those sections of rivers where they have been collected (D. Grout, NHF&G). These areas were scored as 10 (0 - 10 scale) since they in fact must satisfy shad environmental needs (Figure of Juvenile Habitat). Clearly, environmental requirements and tolerances change as juveniles migrate from freshwater nursery areas to the sea.

COMBINED HABITATS

The above spawning, larval and juvenile habitat information were combined into a grid-cell coverage of overall habitat for American shad, using the maximum value from any of the stages on a cell by cell basis (Figure of Combined Life Stages).

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